Studies at St Hilda's College, Oxford; DPhil, Department of Pharmacology, Oxford. Fellow: Department of Physiology, Oxford; College of France, Paris; NYU Medical Center, New York; 1985, Fellow and Tutor in Medicine, Lincoln College; 1996, Professor of Pharmacology, Oxford; since 1988, current position. Honorary Fellow, Royal College of Physicians; 2004, Adelaide Thinker in Residence. Recipient of 29 honorary degrees; Commander of the British Empire; House of Lords, UK; Légion d'Honneur; Fellow, Royal Society of Edinburgh.