Formerly: 27 years at the University of Illinois, Urbana-Champaign, latterly as E.W. and J.M. Gutgsell Endowed Professor of Immunogenetics, with a primary appointment in the Department of Animal Sciences and Member, Center for Advanced Study; served as Director, University of Illinois Biotechnology Center, Founding Director, W.M. Keck Center for Comparative and Functional Genomics, and Founding Director, Institute for Genomic Biology. With the University of California, Davis: 2011-16, Vice-Chancellor for Research; currently, Robert and Rosabel Osborne Endowed Chair and Distinguished Professor of Evolution and Ecology; concurrently, joint appointment in the School of Veterinary Medicine. Current research interest is in mammalian genome evolution in relation to adaptation, speciation, and the origins of cancer. Fellow, American Association for the Advancement of Science. Elected Foreign Member, Royal Swedish Academy of Agriculture and Forestry. Recipient: Wolf Prize in Agriculture (2011); elected to the US National Academy of Sciences (2013).