Graduate, Harvard College and Harvard Law School. Over 30 years' experience in nature conservation. Formerly: Environmental Lawyer, working on environmental protection cases in US; advised UN Environment Programme, co-authored text on environmental law. 1989-2001, with WWF-US, as head of the worldwide conservation programmes and ultimately Executive Vice-President. 2001-05, Director, Conservation and Science programme, David and Lucile Packard Foundation, one of the largest conservation funders in the US. 2005, Director-General, WWF International, serving as Chief Executive Officer, WWF International and, until 2014, leader of the global WWF Network. Since 2014, Consulting Professor, Stanford University.