Graduate, Harvard College and Harvard Law School. Over 30 years' experience in nature conservation. Formerly: Environmental Lawyer, working on environmental protection cases in US; advised UN Environment Programme, co-authored text on environmental law. 1989-2001, with WWF-US, as head of the worldwide conservation programmes and ultimately Executive Vice-President. 2001-05, served as Director, conservation and science programme, David and Lucile Packard Foundation, one of the largest conservation funders in the US. I2005, became Director-General, WWF International, serving as CEO, WWF International and leader of the global WWF Network until 2014. Since 2014, consulting Professor, Stanford University.