1988, MD in Medicine then 1993, MRC in Psychiatry, University of Oxford, UK. 1989-92, Registrar in Psychiatry, Maudsley Hospital, London; with University of Oxford: 1992-95, Wellcome Trust Research Fellow in Mental Health, Institute of Molecular Medicine; 1995-98, Wellcome Trust Career Development Fellow, Institute of Molecular Medicine; 1998-2007, Wellcome Trust Senior Clinical Fellow, then since 2007, Wellcome Trust Principal Fellow, Wellcome Trust Centre for Human Genetics. Wellcome Trust Principal Fellow and Honorary Consultant Psychiatrist; Michael Davys Professor of Neuroscience.